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( g. 12.17). Adenine can pair with adenine if one of the bases undergoes a tautomeric shift while the other rotates about its base-sugar (glycosidic) bond ( g. 12.18). The normal con guration of the base is referred to as the anti con guration; the rotated form is the syn con guration. Since we now believe that as many as 10% of bases

Analysis: We rst determine the equivalent base voltage from equation 96:

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and the equivalent base resistance from equation 97: RB = R1 R2 = 333 k Now we can compute the base current from equation 910: IB = VBB VBE VBB V 5 07 = 128 A = = RB + ( + 1)RE RB + ( + 1)RE 33,000 + 101 3000

base pair can be converted to a TA base pair (a transversion) by way of an intermediate AA base pair. One of the red bases is in the rare tautomeric form, while the other is in the syn con guration. After a second round of DNA replication, one DNA duplex will have a transversion at that point (blue).

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and knowing the current gain of the transistor, , we can determine the collector current: IC = IB = 128 mA Finally, the collector-emitter junction voltage can be computed with reference to equation 911: VCE = VCC IC RC + +1 RE 101 3 10 3 100 = 472 V

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may be in the syn con guration at any moment, the transversion mutagenesis rate should be about 10% of the transition mutagenesis rate, a value not inconsistent with current information. Some base-pair mutations can have serious results. If guanine undergoes an oxidation to 8-oxoguanine ( g. 12.19), it pairs with adenine. A GC base pair is converted to a TA base pair through an 8-oxoguanineadenine intermediate. This transversion has been found to be common in cancers. Since 1953, when Watson and Crick rst described the structure of DNA, tautomerization has been accepted as the obvious source of most transition mutations. However, recent structural data has cast some doubt on this assumption. X-ray crystallography and nuclear magnetic resonance (NMR) studies indicate that both bases in transition mismatches may be in their normal forms. Other mechanisms, similar to wobble base pairing (see chapter 11), may be responsible for most transition mutations. These studies also indicate that some transversions result from direct purine-purine or pyrimidine-pyrimidine base pairing during DNA synthesis. Much work needs to be done to clarify the nature of spontaneous mutagenesis.

= 15 128 10 3 5 10 3 +

Focus on Computer-Aided Tools: An Electronics WorkbenchTM simulation of the circuit analyzed in this example is available in the CD-ROM that accompanies the book

Muller demonstrated that X rays can cause mutation. Certain chemical and temperature treatments can also cause mutation. Determining the mode of action of various chemical mutagens has provided insight into the mutational process as well as the process of carcinogenesis (box 12.1). In addition, knowing how chemical mutagens act has allowed geneticists to produce large numbers of speci c mutations at will (box 12.2).

The material presented in this section has illustrated the basic principles that underlie the operation of a BJT and the determination of its Q point In the next section and later, in 10, we shall develop some simple circuit models for the BJT that will enable us to analyze the transistor ampli er in the linear active region using the familiar tools of linear circuit analysis

95 For the circuit given in Figure 911, the readings are V1 = 3 V, V2 = 24 V, and V3 = 27 V Determine the operating region of the transistor 96 For the circuit given in Figure 921, nd the value of VBB that yields a collector current IC = 63 mA What is the corresponding collector-emitter voltage (assume that VBE = 06 V and that the transistor is in the active region) Assume that RB = 50 k , RE = 200 , RC = 1 k , = 100, and VCC = 14 V 97 What percent change in collector current would result if were changed to 150 in Example 96 Why does the collector current increase by less than 50 percent

N N To deoxyribose N To deoxyribose O Figure 12.18 Transversion mutagenesis. An AA base pair can Figure 12.19 The structure of 8-oxoguanine, which base pairs H N N

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